Inside that custom controller is an Arduino Mega 2560, an LCD, and two 3-axis analog ... on-board µC for autonomous movement as well as wireless, a battery, an IMU, and perhaps some pressure ...

It’s a significant update. The Arduino UNO and Arduino Mega 2560 update the Duemilanove and Mega respectively. They now use an ATMega8U2 instead of an FTDI USB to serial converter chip.